Licensing problem for paper using iNat data

I was under the impression that a CC BY-NC meant you had to give attribution to each user, but you didn’t have to seek permission from them. So it’s possible that listing the contributors in the paper might be sufficient?

https://creativecommons.org/about/cclicenses/

https://creativecommons.org/licenses/by-nc/4.0/

A note about appropriate credit from the cc site: