Web uploader not recognizing scientific binomials or GPS data in photo metadata

In case it helps, here’s the relevant EXIF/IPTC info from a moth photo I just uploaded, where the iNat Uploader successfully got the taxon name (Eudonia philerga) and GPS and description and observation fields. I use darktable instead of Lightroom but they should be tagging in similar ways. I’m not sure which of several similar fields (eg for creation date) iNat is targetting.

File Name : JJS_5519.jpg
File Type : JPEG
Image Description : Attracted in to a 160W Sylania HSB-BW bulb.
GPS Version ID : 2.2.0.0
GPS Altitude Ref : Above Sea Level
Date/Time Original : 2022:10:29 21:38:14
Description : Attracted in to a 160W Sylania HSB-BW bulb.
Subject : Geotagged with iPhone 13 mini-Cyclemeter app, 123A Dyers Pass Road, Eudonia philerga, Insect life stage=adult, Method of surveying for invertebrates=Mercury vapor light sheet, moth light
Hierarchical Subject : Geotagged with iPhone 13 mini-Cyclemeter app, 123A Dyers Pass Road, Eudonia philerga, Insect life stage=adult, Method of surveying for invertebrates=Mercury vapor light sheet, moth light
Create Date : 2022:10:29 21:38:14.50
Date/Time Original : 2022:10:29 21:38:14.500
GPS Altitude : 185.3 m Above Sea Level
GPS Latitude : 43 deg 34’ 46.76" S
GPS Longitude : 172 deg 37’ 59.54" E
GPS Latitude Ref : South
GPS Longitude Ref : East
GPS Position : 43 deg 34’ 46.76" S, 172 deg 37’ 59.54"