This is a second wiki for providing a user-curated list of scientific publications that use iNaturalist data, covering the years 2020 and 2021. We reached the character limit with the first wiki which covers the period up to 2019, and have now reached the character limit with this one too, so have started a third wiki for 2022-2023, and now a fourth for 2024. There should still be space to add any additional older publications to the earlier wikis where appropriate.
Please add relevant publications, including, where possible (1) a link to the paper itself, (2) a link to the observation filter that encompasses as closely as possible what was used by the paper, and (3) contact details for the author(s) in cases where the paper is not open access. There is no need to additionally comment that you have added a paper.
Articles are organised in sections for each year, and then in alphabetical order within year.
These wikis are intended to complement the list of papers that cite GBIF data including iNaturalist exports so papers already on that list need not be included here.
2021
Abercrombie, H. & Hofmann, E. P. (2021) Natural History: Farancia abacura (Red-bellied Mudsnake). Predation. Herpetological Review 52(3): 666. [Observation].
Alexander, P. J. (2021) Pellaea zygophylla, a new combination for a distinctive & well-known but neglected fern. Journal of Semiarid Environments 2: 1â33.
Bastardo, R. H. & Perez-Gelabert, D. E. (2021) Spilostethus pandurus (Hemiptera: Heteroptera: Lygaeidae), nuevo registro para RepĂșblica Dominicana y el Caribe insular. Novitates Caribaea 17: 179â183. [Observation]
Bennett-Smith, M.F., Majoris, J.E., Titus, B.M. & Berumen, M. L. (2021) Clownfish hosting anemones (Anthozoa, Actiniaria) of the Red Sea: new associations and distributions, historical misidentifications, and morphological variability. Marine Biodiversity Records 14: 22. [Forum post]
Braun, H. (2021) An interesting new record of the almost unknown brachypterous phaneropterine katydid Xenicola dohrni (Orthoptera: Tettigoniidae) from Argentina, and its high-ultrasound male-female communication. Zootaxa 4948(2):287â294.
[Observation]
Callaghan, C.T., Poore, A.G., Hofmann, M., Roberts, C.J. & Pereira, H.M. (2021) Large-bodied birds are over-represented in unstructured citizen science data. Scientific Reports 11 (1): 1-11.
DâAngiolella, A. B., Alves, D. S., SodrĂ©, D., Leite, L., Phalan, B., Nascimento, L. R. S. & Diele-Viegas, L. M. (2021) New occurrence records of Lepidodactylus lugubris (DumĂ©ril & Bibron, 1836) (Squamata: Gekkonidae) for the Amazon and Atlantic Forest in Brazil. Cuadernos de HerpetologĂa 35: 189-194.
Delaloye, M., Arregui, M. & Braun, H. (2021) Nuevos registros de la poco conocida Dardarina gaucha Mielke, 1980 (Lepidoptera: Hesperiidae) en la Reserva de la BiĂłsfera Pereyra Iraola,
Buenos Aires, Argentina. Revista Chilena de EntomologĂa 47 (4): 769-774. (https://www.scielo.cl/scielo.php?pid=S0718-89942021000400769&script=sci_abstract)
[Observations]
Douch, J. K. & Poupa, A. M. (2021) Citizen science data opens multiple avenues for iridovirus research and prompts first detection of Invertebrate iridescent virus 31 in Australia. Journal of Invertebrate Pathology 183: 107619.
Durso, A. M., I. Bolon, A. R. Kleinhesselink, M. R. Mondardini, J. L. Fernandez-Marquez, F. Gutsche-Jones, C. Gwilliams, M. Tanner, C. E. Smith, W. WĂŒster, F. Grey, & R. Ruiz de Castañeda (2021) Crowdsourcing snake identification with online communities of professional herpetologists and avocational snake enthusiasts. Royal Society Open Science 8: 201273.
Eiseman, C., O. Lonsdale, J. Van Der Linden, T. Feldman, M. W. Palmer (2021) Thirteen new species of Agromyzidae (Diptera) from the United States, with new host and distribution records for 32 additional species. Zootaxa 4931: 1â68. (Contact Authorâs iNat account, @ ceiseman)
Forister, M. L., C. A. Halsch, C. C. Nice, J. A. Fordyce, T. E. Dilts, J. C. Oliver, K. L. Prudic, A. M. Shapiro, J. K. Wilson & J. Glassberg (2021) Fewer butterflies seen by community scientists across the warming and drying landscapes of the American West. Science 371: 1042â1045.
Friesen, C. & Zhang, Y. M. (2021) Rose gall wasps (Cynipidae: Diplolepis) of Manitoba, including a new provincial record. Proceedings of the Entomological Society of Manitoba. 77: 52â60.
Grosso-Silva, J. M., da Silva, V., Silva, Ă. & van der Heyden, T. (2021) Sastrapada Amyot & Serville, 1843 (Hemiptera: Reduviidae), new genus for Portugal. Arquivos EntomolĂłxicos 24: 347â350.
Guariento, E. & Franzini, G. (2021) Attalus minimus (Insecta, Melyridae) recently rediscovered in South Tyrol after 109 years. Gredleriana 21: 169â171. Observations used by the paper
Hausdorf, B., Parr, M., Shappell, L. J., Oldeland, J. & Robinson, D. G. (2021) The introduction of the European Caucasotachea vindobonensis (Gastropoda: Helicidae) in North America, its origin and its potential range. Biological Invasions https://doi.org/10.1007/s10530-021-02579-4
Heron, K. M. D. Windham, D. R. Farrar & K. M. Pryer (2021) Looking Back on 130 Years of Fern and Lycophyte Research in Glacier National Park, Montana: A Modern Taxonomic Account. American Fern Journal 111(4), 223-250, (5 November 2021). https://doi.org/10.1640/0002-8444-111.4.223. [author: heron.keegan@gmail.com]
Hitchcock, C., Sullivan, J. and OâDonnell, K., 2021. Cultivating Bioliteracy, Biodiscovery, Data Literacy, and Ecological Monitoring in Undergraduate Courses with iNaturalist. Citizen Science: Theory and Practice , 6(1), p.26. [DOI: http://doi.org/10.5334/cstp.439](DOI: http://doi.org/10.5334/cstp.439)
Hofmann, E.P. & S.D. Wilkinson (2021) Attempted predation of a black swampsnake (Liodytes pygaea) by a Glossy Ibis (Plegadis falcinellus) in Orange County, Florida. Florida Field Naturalist 49(4): 158â161. [Observation]
Kolby, J.E. (2021) Identification Guide to Tortoises and Freshwater Turtles: Parts, Products and Derivatives in Trade Commissioned by CITES Secretariat, Geneva, Switzerland.
Leaché AD, Davis HR, Singhal S, Fujita MK, Lahti ME and Zamudio KR (2021) Phylogenomic Assessment of Biodiversity Using a Reference-Based Taxonomy: An Example With Horned Lizards (Phrynosoma ). Front. Ecol. Evol. 9:678110.
Mesaglio, T. & Callaghan, C.T. (2021) An overview of the history, current contributions and future outlook of iNaturalist in Australia. Wildlife Research 48(4), 289-303.
Mesaglio, T., Soh, A., Kurniawidjaja, S. & Sexton, C. (2021) âFirst Known Photographs of Living Specimensâ: the power of iNaturalist for recording rare tropical butterflies. Journal of Insect Conservation 25(5), 905-911.
[based on https://www.inaturalist.org/projects/first-known-photographs-of-living-specimens]
Neill, A.K. & S.R. Kieschnick (2021) Noccaea perfoliata or Microthlaspi perfoliatum (Brassicaceae), new to the flora of Texas, U.S.A. J. Bot. Res. Inst. Texas 15(1).
Nelson, T.D., & C.E. Moffat. (2021) A northern range extension of a Canadian species of special concern, Dielis pilipes (Hymenoptera: Scoliidae), in the Okanagan Valley of British Columbia. Journal of the Entomological Society of British Columbia. 118: 82â86.
Pintar, M.R., J.L. Kline, & J.C. Trexler. (2021). The aquatic Heteroptera (Hemiptera) of marshes in the Florida Everglades. Florida Entomologist 104(4): 308â319.
Putman, B. J., R. Williams, E. Li, & G. B. Pauly (2021) The power of community science to quantify ecological interactions in cities. Scientific Reports 11: 3069.
Schneider, A.C. & B.E. Benton (2021) Morphometrics and redescription of Aphyllon fasciculatum and Aphyllon franciscanum, two widespread but previously conflated species in western North America. Systematic Botany 46(2): 446â455. (observations used would be a small subset of this search)
Seidel, M., LĂŒttke, M., Cocquempot, C., Potts, K., Heeney, W.J. & Husemann, M. (2021) Citizen scientists significantly improve our knowledge on the non-native longhorn beetle Chlorophorus annularis (Fabricius, 1787) (Coleoptera, Cerambycidae) in Europe. BioRisk 16: 1-13. [original iNat observation 1; original iNat observation 2]
Sexton, C. (2021). Identification and distribution of the Petrophila fulicalis species group (Crambidae): Taking advantage of citizen science data. J. Lepidopteristsâ Society 75(2):113-127. Also available on researchgate.net.
Sexton, C. (2021). Fire, Rain, and Butterflies in the Chisos Mountains of Texas Southern Lepidopteristsâ News 43(4):335-342.
Sexton, C. (2021). Bioblitz at Maddin Prairie, Mitchell Co., Texas, August 21-22, 2021 Southern Lepidopteristsâ News 43(4):349-350.
Shirey, V., Belitz, M.W., Barve, V. & R. Guralnick (2021) A complete inventory of North American butterfly occurrence data: Narrowing data gaps, but increasing bias. Ecography 44(4): 537-547.
Soteropoulos, D.L., De Bellis, C.R. & Witsell, T. (2021) Citizen science contributions to address biodiversity loss and conservation planning in a rapidly developing region. Diversity 13(6): 255.
van der Heyden, T. (2021) First records of Leptoglossus occidentalis Heidemann, 1910 (Hemiptera: Heteroptera: Coreidae) in Estonia and Belarus. Heteroptera Poloniae â Acta Faunistica 15: 5â6.
van der Heyden, T. (2021) Erstfund von Zelus renardii Kolenati, 1856 in Deutschland (Heteroptera: Reduviidae). Heteropteron 61: 31â32.
van der Heyden, T. (2021) On the recent Northern European dispersion of Zelus renardii Kolenati (Hemiptera: Heteroptera: Reduviidae) via human activity. Israel Journal of Entomology 51: 43â46.
van der Heyden, T. (2021) Thaumastocoris peregrinus Carpintero & DellapĂ©, 2006 (Hemiptera: Heteroptera: Thaumastocoridae) has reached the Greek island of Crete. Journal of the Heteroptera of Turkey 3(1): 11â13.
van der Heyden, T. (2021) First records of Spilostethus furcula (Herrich-SchĂ€ffer, 1850) (Hemiptera: Heteroptera: Lygaeidae) in Malta. Revista gaditana de EntomologĂa XII(1): 127â129.
van der Heyden, T. (2021) First records of Hierodula transcaucasica Brunner von Wattenwyl, 1878 in Slovenia and Spain (Mantodea: Mantidae). Arquivos EntomolĂłxicos 24: 265â266.
van der Heyden, T. (2021) Confirmation of the presence ofSolenosthedium bilunatum (LefĂšbvre, 1827) (Hemiptera: Heteroptera: Scutelleridae) in Albania. Journal of the Heteroptera of Turkey 3(2): 111â113.
van der Heyden, T. & Schwarz, C. J. (2021) New data on the presence of Hierodula transcaucasica Brunner von Wattenwyl, 1878 on Crete (Mantodea: Mantidae). BV news Publicaciones CientĂficas 10(118): 6â12.
Van Eupen, C., D. Maes, M. Herremans, K. R. R. Swinnen, B. Somers & S. Luca. (2021) The impact of data quality filtering of opportunistic citizen science data on species distribution model performance.. Ecological Modelling 444:109453.
2020
Alexander, P.J. (2020) Muilla lordsburgana (Asparagaceae: Brodiaeoideae), a new species found north of Lordsburg, southwestern New Mexico. Journal of Semiarid Environments 1: 1-10.
Barve, V., L. Brenskelle, D., Daijiang, B. Stucky, N. Barve, M. Hantak, B. McLean, D. Paluh, J. Oswald, M. Belitz, R. A. Folk & R. P. Guralnick (2020) Methods for broad-scale plant phenology assessments using citizen scientistsâ photographs. Applications in Plant Sciences 8 (1): e11315
Callaghan, C.T., Poore, A.G., Mesaglio, T., Moles, A.T., Nakagawa, S., Roberts, C., Rowley, J.J., Verges, A., Wilshire, J.H. & Cornwell, W.K. (2020) Three frontiers for the future of biodiversity research using citizen science data. BioScience 71: 55â63.
Chatfield-Taylor, W. (2020) Predator avoidance leads to separate emergence cycles in the protoperiodical Okanagana magnifica Davis, 1919 (Hemiptera: Cicadidae). Pan-Pacific Entomologist 96(3): 1â7
Chernov, E., E.P. Hofmann & W. WĂŒster (2020) Scolecophis atrocinctus (Black-banded Snake). Reproduction/combat behavior. Herpetological Review 51(3): 628â629. [original iNaturalist observation]
Chowdhury, S., M. F. Braby & R. A. Fuller (2020) Coasting along to a wider range: niche conservatism in the recent range expansion of the Tawny Coster, Acraea terpsicore (Lepidoptera: Nymphalidae). Diversity and Distributions 27(3): 402â415. [Observations]
Crews, S. C., W. Galvis & L. A. Esposito (2020) Gliding canopy flatties and relatives from the Selenops banksi group (Araneae: Selenopidae). Journal of Natural History 54(35â36): 2343â2365.
Cutler, B. & M. Parr. (2020) First records of Icius subinermis (Araneae: Salticidae) in North America, with notes on the local establishment of this species and its behavior in captivity. Peckhamia 226.1: 1â5.
DĂaz-Calafat, J. (2020). First record of Sceliphron caementarium (Drury, 1773) (Hymenoptera: Sphecidae) in the Balearic Islands, with an identification key to the archipelagoâs species of the genus Sceliphron Klug, 1801. Ecosistemas 29(1):1939. [original observation].
Esparza, J. E. & G. B. Pauly (2020) Elgaria multicarinata (Southern Alligator Lizard). High elevation record. Herpetological Review 51: 124. [observation] [additional iNaturalist observations are mentioned in the publication]
Hernandez, Jason P. (2020) Scientific Note: Observations on the life history of Hyalurga vinosa (Lepidoptera: Erebidae). Tropical Lepidoptera Research 30(2): 90â92.
Hernawati, D., Chaidir, D. M. & Meylani, V. (2020) The use of iNaturalist on learning courses of zoology vertebrates for prospective biology teachers. Journal of Physics: Conference Series 1440(1): 012064.
Hewitt, S. J., Atha, D. & Ringer, D. J. (2020) Descurainia sophia (L.) Webb ex Prantl (Brassicaceae) rediscovered in New York City after more than 100 years. New York Flora Association Quarterly Newsletter (Spring 2020) 31(2): 10â12.
Hochmair, H. H., Scheffrahn, R. H., Basille, M. & Boone, M. (2020) Evaluating the data quality of iNaturalist termite records. PLoS ONE 15 (5): e0226534.
Kirchhoff, C., Callaghan, C.T., Keith, D.A., Indiarto, D., Taseski, G., Ooi, M.K., Le Breton, T.D., Mesaglio, T., Kingsford, R.T. & Cornwell, W.K. (2020) Rapidly mapping fire effects on biodiversity at a large-scale using citizen science. Science of The Total Environment 755: 142348.
Klee, B. (2020) Summer insect pandemics in the United States. Wolfram Demonstrations Project.
Lanner, J., K. Huchler, B. Pachinger, C. Sedivy, & H. Meimberg (2020) Dispersal patterns of an introduced wild bee, Megachile sculpturalis Smith, 1853 (Hymenoptera: Megachilidae) in European alpine countries. PLoS ONE 15(7): e0236042.
Lebel, T., Syme, K., Barrett, M. D. & Cooper, J. A. (2002) Two new species of Asproinocybe (Tricholomataceae) from Australia. Muelleria 38: 77â85.
Lehtinen, R. M., Carlson, B. M., Hamm, A. R., Riley, A. G., Mullin, M. M., & Gray, W. J. (2020) Dispatches from the neighborhood watch: Using citizen science and field survey data to document color morph frequency in space and time. Ecology and Evolution 10(3): 1526â1538.
Lupoli, R., van der Heyden, T. & Dioli, P. (2020) Erthesina Spinola, 1837 â a new alien genus for Europe found in Albania (Hemiptera: Pentatomidae). Heteroptera Poloniae â Acta Faunistica 14: 121â123.
Oficialdegui, Francisco J., Sanchez, Marta I., Clavero, Miguel (2020) One century away from home: how the red swamp crayfish took over the world. Reviews in Fish Biology and Fisheries 30: 121â135.
Pauly, G. B., Shaulsky, M. C., Barley, A. J., Kennedy-Gold, S., Stewart, S. C., Keeny, S. & Thomson, R. C. (2020) Morphological change during rapid population expansion confounds Leopard Frog identifications in the Southwestern United States Copeia 108(2): 299â308.
Pintar, MR, and JL Keller. 2020. First record of Hydrophilus ensifer Brulle (Coleoptera: Hydrophilidae) in the continental United States. The Coleopterists Bulletin 74 (4): 696-698.
Reith, M. (2020) Rediscovery of Gesneria brachysepala Urb. & Ekman. Gesneriads, the Journal for Gesneriad Growers 70(1) [Observation]
Reyes, F. O., Landestoy T., M. A., & JimĂ©nez, L., (2020). Geographic distribution: Gymnophthalmus underwoodi (b) Herpetological Review 51 (3): 539â540. [Observation]
Sexton, C. (2020). Fingerprinting moths in the field: An example with the Texas endemic âCanyonlands Quakerâ, Tricholita baranca* (Noctuidae) Southern Lepidopteristsâ News 42(4):309-311. [Observations]
Skejo, J., Connors, M., Hendriksen, M., Lambert, N., Chong, G., McMaster, I., Monaghan, N., Rentz, D., Richter, R., Rose, K. & FranjeviÄ, D. (2020) Online social media tells a story of Anaselina, Paraselina, and Selivinga (Orthoptera, Tetrigidae), rare Australian pygmy grasshoppers. ZooKeys 948: 107â119.
Teofilova, Teodora & Naumova, Maria. (2020). First record of Lilioceris faldermanni (GuĂ©rin-MĂ©neville, 1829) (Coleoptera: Chrysomelidae: Criocerinae) in Bulgaria, including a record of a new host plant. Acta Zoologica Bulgarica. 72. 27â30. [obs. in Fig. 1:4 & Fig. 2:f; obs. in Fig. 1:5] [Note that records are not directly cited, but the photos in Fig. 2 clearly come from the observation and the observer confirms their records were used in this paper]
Uyeda, K.A., Stow, D.A. & Richart, C.H. Assessment of volunteered geographic information for vegetation mapping. Environ Monit Assess 192, 554 (2020). https://doi.org/10.1007/s10661-020-08522-9 , Formatted read-only version, Authorâs Approved Manuscript, Observations
van der Heyden, T. (2020) First record of Leptoglossus occidentalis Heidemann, 1910 in Guatemala (Hemiptera, Heteroptera, Coreidae). BoletĂn de la AsociaciĂłn Española de EntomologĂa 44(1-2): 213â214.
van der Heyden, T. (2020) First record of Corythauma ayyari (Drake, 1933) (Hemiptera: Heteroptera: Tingidae) in Egypt. Journal of the Heteroptera of Turkey 2(1): 1â2.
van der Heyden, T. (2020) First records of Leptoglossus talamancanus Brailovsky & Barrera, 1998 (Hemiptera: Heteroptera: Coreidae) for Belize and Mexico. BoletĂn del Museo Nacional de Historia Natural del Paraguay 24(1): 45â46.
van der Heyden, T. (2020) Records of Solenosthedium bilunatum (LefĂšbvre, 1827) on the Italian island of Ustica and the Spanish island of Ibiza (Hemiptera: Heteroptera: Scutelleridae). Arquivos EntomolĂłxicos 22: 289â291.
van der Heyden, T. (2020) First record of Leptoglossus cartagoensis Brailovsky & Barrera, 1998 (Hemiptera: Heteroptera: Coreidae) for Panama. BoletĂn del Museo Nacional de Historia Natural del Paraguay 24(2): 55â56.
van der Heyden, T. (2020) Confirmation of the presence of Spilostethus furcula (Herrich-SchĂ€ffer, 1850) (Heteroptera: Lygaeidae) on the Balearic Islands (Spain). Revista gaditana de EntomologĂa XI: 133â135.
van der Heyden, T. (2020) First record of Leptoglossus occidentalis Heidemann, 1910 (Hemiptera: Coreidae) in Andorra. Arquivos EntomolĂłxicos 22: 377â378.
van der Heyden, T. (2020) First record of Sphedanolestes cingulatus (Fieber, 1864) (Hemiptera: Heteroptera: Reduviidae) in Greece. Journal of the Heteroptera of Turkey 2(2): 148â149.
van der Heyden, T. & Dioli, P. (2020) First records of Lasiocoris crassicornis (Lucas, 1849) (Heteroptera: Rhyparochromidae) in Lebanon and Israel. Heteroptera Poloniae â Acta Faunistica 14: 119â120.
van der Heyden, T. & FaĂșndez, E. I. (2020) First records of Leptoglossus occidentalis Heidemann, 1910 (Hemiptera:Heteroptera: Coreidae) in Brazil and South Africa. BoletĂn del Museo Nacional de Historia Natural del Paraguay 24(1): 28â30.
van der Heyden, T. & Grosso-Silva, J. M. (2020) First record of Zelus renardii Kolenati, 1856 in Portugal (Heteroptera: Reduviidae: Harpactorinae). Arquivos EntomolĂłxicos 22: 347â349.
Vink, C. & Curtis, K. M. (2020). A redescription of Philoponella congregabilis, an Australian hackled orb weaver spider (Uloboridae) now found in Christchurch, New Zealand. Records of the Canterbury Museum 34: 85â94.
Werenkraut, V., Baudino, F., & Roy, H. E. (2020). Citizen science reveals the distribution of the invasive harlequin ladybird (Harmonia axyridis Pallas) in Argentina. Biological Invasions 22: 2915â2921.
White, E.L., M. D. Schlesinger, and T.G. Howard. 2022. The Empire State
Native Pollinator Survey (2017-2021). New York Natural Heritage Program, Albany, NY.
Wilson, Joseph S.; Pan, Aaron D.; General, David Emmanual M.; Koch, Jonathan B. (2020) More eyes on the prize: an observation of a very rare, threatened species of Philippine Bumble bee, Bombus irisanensis, on iNaturalist and the importance of citizen science in conservation biology. Journal of Insect Conservation 24: 727â729 [observation, observation]
Woznica J. A. (2020): Pseudoleria pectinata (Loew, 1872) â a new genus and species of heleomyzid fly introduced into the European fauna (Diptera: Heleomyzidae). Acta Zoologica Cracoviensia 63(2): 23â27 [original observation]
